In the context of the anime, this statement refers to the Orochi, which is the enemy of human beings.

However, based on the text alone, what species of animal comes to mind?

Donten ni Warau Episode 6
donten 10 donten 11

greed, avarice, pride“: common traits of human beings
devouring every living thing“: humans consume animals, plants and fungi
burning the land“: humans slash-and-burn to clear forests
drying the seas“: humans build dams in rivers
destruction“: pollution by humans has ruined the natural landscape

Kiseijuu: Sei no Kakuritsu Episode 8
kiseijuu 33

Therefore, who is the true enemy of human beings?

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